diff --git a/platform/android/SCsub b/platform/android/SCsub
index bbe2c1b6791..f97ad7a8a70 100644
--- a/platform/android/SCsub
+++ b/platform/android/SCsub
@@ -97,12 +97,21 @@ if lib_arch_dir != "" and host_subpath != "":
     else:
         lib_tools_dir = ""
 
-    out_dir = "#platform/android/java/lib/libs/" + lib_tools_dir + lib_type_dir + "/" + lib_arch_dir
+    jni_libs_dir = "#platform/android/java/lib/libs/" + lib_tools_dir + lib_type_dir + "/"
+    out_dir = jni_libs_dir + lib_arch_dir
     env_android.CommandNoCache(out_dir + "/libgodot_android.so", lib, Move("$TARGET", "$SOURCE"))
 
     stl_lib_path = f"{env['ANDROID_NDK_ROOT']}/toolchains/llvm/prebuilt/{host_subpath}/sysroot/usr/lib/{triple_target_dir}/libc++_shared.so"
     env_android.CommandNoCache(out_dir + "/libc++_shared.so", stl_lib_path, Copy("$TARGET", "$SOURCE"))
 
+    if env["debug_symbols"] and env["separate_debug_symbols"]:
+        debug_symbols_zip_file = (
+            ("#bin/android-editor-" if env.editor_build else "#bin/android-template-")
+            + lib_type_dir
+            + "-native-symbols.zip"
+        )
+        env_android.NoCache(Zip(debug_symbols_zip_file, jni_libs_dir, ZIPROOT=Dir(jni_libs_dir)))
+
     if env["generate_android_binaries"]:
         env_android.AlwaysBuild(
             env_android.CommandNoCache(
diff --git a/platform/android/java/app/build.gradle b/platform/android/java/app/build.gradle
index 658144eaa2f..b13a42633b6 100644
--- a/platform/android/java/app/build.gradle
+++ b/platform/android/java/app/build.gradle
@@ -101,7 +101,7 @@ android {
         }
 
         ndk {
-            debugSymbolLevel 'FULL'
+            debugSymbolLevel 'NONE'
             String[] export_abi_list = getExportEnabledABIs()
             abiFilters export_abi_list
         }
@@ -127,9 +127,11 @@ android {
         exclude 'META-INF/LICENSE'
         exclude 'META-INF/NOTICE'
 
-        // 'doNotStrip' is enabled for development within Android Studio
+        // Debug symbols are kept for development within Android Studio.
         if (shouldNotStrip()) {
-            doNotStrip '**/*.so'
+            jniLibs {
+                keepDebugSymbols += '**/*.so'
+            }
         }
 
         // Always select Godot's version of libc++_shared.so in case deps have their own
diff --git a/platform/android/java/build.gradle b/platform/android/java/build.gradle
index 94433c7ab5d..96aec0188db 100644
--- a/platform/android/java/build.gradle
+++ b/platform/android/java/build.gradle
@@ -176,10 +176,6 @@ def generateBuildTasks(String flavor = "template", String edition = "standard",
                         from("app/build/outputs/apk/${edition}/${target}") {
                             include("android_${filenameSuffix}.apk")
                         }
-                        from("app/build/outputs/native-debug-symbols/${edition}${capitalizedTarget}") {
-                            include("native-debug-symbols.zip")
-                            rename ("native-debug-symbols.zip", "android-template-${edition}-${target}-native-debug-symbols.zip")
-                        }
                         into(binDir)
                     }
                 }
@@ -194,10 +190,6 @@ def generateBuildTasks(String flavor = "template", String edition = "standard",
                         from("editor/build/outputs/apk/${androidDistro}/${target}") {
                             include("android_editor-${androidDistro}-${target}*.apk")
                         }
-                        from("editor/build/outputs/native-debug-symbols/${androidDistro}${capitalizedTarget}") {
-                            include("native-debug-symbols.zip")
-                            rename ("native-debug-symbols.zip", "android-editor-${androidDistro}-${target}-native-debug-symbols.zip")
-                        }
                         into(androidEditorBuildsDir)
                     }
                 }
@@ -323,17 +315,11 @@ task cleanGodotTemplates(type: Delete) {
 
     // Delete the Godot templates in the Godot bin directory
     delete("$binDir/android_debug.apk")
-    delete("$binDir/android-template-standard-debug-native-debug-symbols.zip")
     delete("$binDir/android_dev.apk")
-    delete("$binDir/android-template-standard-dev-native-debug-symbols.zip")
     delete("$binDir/android_release.apk")
-    delete("$binDir/android-template-standard-release-native-debug-symbols.zip")
     delete("$binDir/android_monoDebug.apk")
-    delete("$binDir/android-template-mono-debug-native-debug-symbols.zip")
     delete("$binDir/android_monoDev.apk")
-    delete("$binDir/android-template-mono-dev-native-debug-symbols.zip")
     delete("$binDir/android_monoRelease.apk")
-    delete("$binDir/android-template-mono-release-native-debug-symbols.zip")
     delete("$binDir/android_source.zip")
     delete("$binDir/godot-lib.template_debug.aar")
     delete("$binDir/godot-lib.template_debug.dev.aar")
@@ -343,4 +329,12 @@ task cleanGodotTemplates(type: Delete) {
     delete("$binDir/godot-lib.debug.aar")
     delete("$binDir/godot-lib.dev.aar")
     delete("$binDir/godot-lib.release.aar")
+
+    // Delete the native debug symbols files.
+    delete("$binDir/android-editor-debug-native-symbols.zip")
+    delete("$binDir/android-editor-dev-native-symbols.zip")
+    delete("$binDir/android-editor-release-native-symbols.zip")
+    delete("$binDir/android-template-debug-native-symbols.zip")
+    delete("$binDir/android-template-dev-native-symbols.zip")
+    delete("$binDir/android-template-release-native-symbols.zip")
 }
diff --git a/platform/android/java/editor/build.gradle b/platform/android/java/editor/build.gradle
index 82c6ad02e59..ef846b270cd 100644
--- a/platform/android/java/editor/build.gradle
+++ b/platform/android/java/editor/build.gradle
@@ -90,7 +90,7 @@ android {
             editorBuildSuffix: ""
         ]
 
-        ndk { debugSymbolLevel 'FULL' }
+        ndk { debugSymbolLevel 'NONE' }
     }
 
     base {
@@ -141,9 +141,11 @@ android {
     }
 
     packagingOptions {
-        // 'doNotStrip' is enabled for development within Android Studio
+        // Debug symbols are kept for development within Android Studio.
         if (shouldNotStrip()) {
-            doNotStrip '**/*.so'
+            jniLibs {
+                keepDebugSymbols += '**/*.so'
+            }
         }
     }
 
diff --git a/platform/android/java/lib/build.gradle b/platform/android/java/lib/build.gradle
index a3ff1fa9dd0..4e8ff5041ec 100644
--- a/platform/android/java/lib/build.gradle
+++ b/platform/android/java/lib/build.gradle
@@ -67,9 +67,11 @@ android {
         exclude 'META-INF/LICENSE'
         exclude 'META-INF/NOTICE'
 
-        // 'doNotStrip' is enabled for development within Android Studio
+        // Debug symbols are kept for development within Android Studio.
         if (shouldNotStrip()) {
-            doNotStrip '**/*.so'
+            jniLibs {
+                keepDebugSymbols += '**/*.so'
+            }
         }
     }
 
diff --git a/platform/android/platform_android_builders.py b/platform/android/platform_android_builders.py
index 4593e8a8058..dbd70dc9f0c 100644
--- a/platform/android/platform_android_builders.py
+++ b/platform/android/platform_android_builders.py
@@ -19,7 +19,10 @@ def generate_android_binaries(target, source, env):
     if env["target"] == "editor":
         gradle_process += ["generateGodotEditor", "generateGodotHorizonOSEditor", "generateGodotPicoOSEditor"]
     else:
-        gradle_process += ["generateGodotTemplates"]
+        if env["module_mono_enabled"]:
+            gradle_process += ["generateGodotMonoTemplates"]
+        else:
+            gradle_process += ["generateGodotTemplates"]
     gradle_process += ["--quiet"]
 
     if env["debug_symbols"] and not env["separate_debug_symbols"]: